DirectStream, a.k.a. UPnP, is a feature which enables Orb to send streams directly from your Orb PC to your Orb client device (i.e., other PC or PDA) without going through Orb's stream servers. This feature is automatically activated by Orb if Orb detects that you have a router with the UPnP functionality enabled.
If UPnP is enabled on your router but the Use UPnP for DirectStream checkbox is greyed out in your Orb tray icon (Configuration > Advanced tab), Orb cannot detect the UPnP feature on your router, possibly due to an older firmware version. You may want to upgrade your router's firmware.
Note: within your LAN (Local Area Network), Orb will always stream directly from the Orb server to the Orb client if it detects that both your PC and your client device share the same public IP address. You should see a significant improvement when using DirectStream on your local network.